垫状卷柏Selaginella pulvinata (Hook. et Grev.) Maxim
垫状卷柏Selaginella pulvinata (Hook. et Grev.) Maxim
7.垫状卷柏（中国主要植物图说·蕨类植物门）图版23：1-8
Selaginella pulvinata (Hook. et Grev.) Maxim. in Mem. Acad. Imp. Sci. Petersb. 9: 335. 1859; Hand. -Mazz., Symb. Sin. 6: 5. 1929; Reed, Index Selaginellarum, inMem. Soc. Brot. 18: 187. 1966; Ic. Corm. Sin. 1: 111. 1972; Fl. Fujian. 1: 11. 1982; Fl. Xizang. 1: 24. 1983: B. D. Liu et al. in Bull. Bot. Res. (Harbin) 9 (3): 114, Pl. 5: 1-2. 1989; P. S. Wang in J. Arnold Arbor. 71: 265. 1990: R. D. Dixit, Selag. India 51, f. 16, Pl. 17. 1992; Fl. Jiangxi 1: 31, f. 16. 1993; Vasc. PI. Heng-duan Mts. 1: 9. 1993: P. S. Wang et X. Y. Wang, Pterid. Fl. Guizhou 639, pl. 155, f. 11-12. 2001. ——Lycopodium pulvinatum Hook. et Grev. in Hook. Bot. Misc. 2: 381. 1831. ——S. tamariscina var. pulvinata (Hook. et Grev.) Alston in Bull. FanMem. Inst. Biol. 5: 271. 1934, et in Proc. Nat. Inst. Sci. India 11: 221. 1945; S. H. Fu, Ill. Important Chinese Pl., Pterid. 8. 1957; Reed, Index Selaginellarum, in Mem.Soc. Brot. 18: 223. 1966; Fl. Henan 1: 5. 1981: R. D. Dixit, Cens. Indian Pterid. 16. 1984; Fl. Intramong. 1: 55, Pl. 2, f. 18. 1985: Fl. Liaoning. 1: 15. 1988; J. z. Wang et S. X. Li, Clav. Pl. Chin. Bor. -Ori. 2nd ed. 22. 1995. ——Lycopodioides pulv-inata (Hook. et Grev.) H. S. Kung, Fl. Sichuan. 6: 64, pl. 18, f. 1-3. 1988. ——S. involvens auct. non (Sw.) Spring: Baker, Handb. Fern-Allies 87, no. 204. 1887, p. p. : Warb. in Monsunia 1: 113. 1900, P. P. quoad pl. Peking; O. C. Schmidt in ActaHorti Gothob. 5: 52. 1930. ——S. tamariscina auct. non (P. Beauv.) Spring in Alstonin Lecomte, Fl. Gen. Indo-Chine 7 (2): 559. 1951; P. H. Ho, Ill. Fl. Vietnam 1: 27, cum fig. 1991.
土生或石生，旱生复苏植物，呈垫状，无匍匐根状茎或游走茎。根托只生于茎的基部，长2-4厘米，直径0.2-0.4毫米，根多分叉，密被毛，和茎及分枝密集形成树状主干，高数厘米。主茎自近基部羽状分枝，不呈“之”字形，禾秆色或棕色，主茎下部直径1毫米，不具沟槽，光滑，维管束1条； 侧枝4-7对，2-3回羽状分枝，小枝排列紧密，主茎上相邻分枝相距约1厘米，分枝无毛，背腹压扁，主茎在分枝部分中部连叶宽2.2-2.4毫米，末回分枝连叶宽1.2-1.6毫米。叶全部交互排列，二形，叶质厚，表面光滑，不具白边，主茎上的叶略大于分枝上的叶，相互重叠，绿色或棕色，斜升，边缘撕裂状。分枝上的腋叶对称，卵圆形到三角形，2.5毫米X 1.0毫米，边缘撕裂状并具睫毛。小枝上的叶斜卵形或三角形，2.8-3.1毫米XO.9-1.2毫米，覆瓦状排列，背部不呈龙骨状，先端具芒，基部平截（具簇毛），边缘撕裂状，并外卷。侧叶不对称，小枝上的叶距圆形，略斜升，2.9-3.2毫米X1.4-1.5毫米，先端具芒，边缘全缘，基部上侧扩大，加宽，覆盖小枝，基部上侧边缘不为全缘，呈撕裂状，基部下侧不呈耳状，边缘不为全缘，呈撕裂状，下侧边缘内卷。孢子叶穗紧密，四棱柱形，单生于小枝末端，10-20毫米X 1.5-2.0毫米；孢子叶一形，不具白边，边缘撕裂状，具睫毛；大孢子叶分布于孢子叶穗下部的下侧或中部的下侧或上部的下侧。大孢子黄白色或深褐色；小孢子浅黄色。
产山西、北京、重庆（城口、巫溪、秀山）、福建（德化、南靖）、甘肃（宕昌、文县）、广西（德宝、隆林）、贵州（册亨、从江、大方、关岭、贵阳、赫章、晴隆、黔西、水城、松桃、铜仁、威宁、西秀、兴义、镇宁）、河北（阜平、临城、山海关、蔚县、行唐、邢台、元氏、琢鹿）、河南（内乡）、江西（赣北）、辽宁（凌源）、陕西（眉县）、四川（安远、宝兴、北川、布拖、丹巴、道孚、德昌、都江堰、峨边、甘洛、高县、汉源、黑水、会理、剑阁、金川、金阳、九龙、康定、雷波、里塘、理县、芦山、泸定、马尔康、茂县、米易、冕宁、木里、南坪、宁南、平武、青川、石棉、汶川、西昌、喜德、乡里、小金、新龙、雅江、盐边、越西）、台湾、西藏（昌都、甘孜、拉萨、朗县、林周、芒康、米林、盐井）、云南（德钦、贡山、昆明、中甸）。常见于石灰岩上，海拔（100-) 1000-3000 (-4250)米。也分布到蒙古、俄罗斯西伯利亚、朝鲜半岛、日本、印度北部、越南、泰国等地。模式标本采自印度。
《Flora of China》 Vol. 2-3 (2013)
Selaginella pulvinata (Hooker & Greville) Maximowicz Mém. Acad. Imp. Sci. Saint Pétersbourg, sér. 7. 9: 335. 1859.
垫状卷柏 dian zhuang juan bai
Lycopodium pulvinatum Hooker & Greville, Hooker’s J. Bot. Kew Gard. Misc. 2: 381. 1831; Lycopodioides pulvinata (Hooker & Greville) H. S. Kung; Selaginella tamariscina (P. Beauvois) Spring var. pulvinata (Hooker & Greville) Alston.
Plants terrestrial or epilithic, xerophytic, “resurrectional”; stems forming rosette, seasonally green, stems and roots entangled forming treelike trunk, plants 2-10 cm, without creeping rhizomes. Rhizophores restricted to base of stem, 2-4 cm, rather thick or slender, much forked at end and forming thick massive rootstock. Main stems branched near and above base, pinnately branched, stramineous or brown, main stem ca. 1 mm in diam. at lower part; primary leafy branches 4-7 pairs, 2 or 3 times pinnately branched, branchlets dense, adjacent primary branches on main stem ca. 1 cm apart, leafy portion of main stem including leaves 2.2-2.4 mm wide at middle, ultimate branches 1.2-1.6 mm wide including leaves. Axillary leaves on main stems larger than those on branches; axillary leaves on branches symmetrical, ovate to triangular, ca. 2.5 × 1 mm, base exauriculate, margin lacerate-ciliolate. Dorsal leaves on branches imbricate, obliquely ovate or triangular, 2.8-3.1 × 0.9-1.2 mm, not carinate, base truncate (with a tuft of hairs), margin lacerate, outer margin revolute, apex aristate. Ventral leaves slightly ascending, asymmetrical, oblong, 2.9-3.2 × 1.4-1.5 mm, apex aristate, entire; basiscopic base not auriculate, margin lacerate, lower margin involute; acroscopic base enlarged, broader, overlapping stem and branches, margin lacerate. Strobili solitary, terminal, compact, tetragonal, 10-20 × 1.5-2 mm; sporophylls uniform, not white-margined, margin lacerate-ciliolate; megasporophylls in basal portion of strobili on lower side, in middle on lower side, or in upper portion on lower side; microspores pale yellow, megaspores white-yellow or dark brown.
Common in rock crevices on exposed rocky (limestone) ridges; (100-)1000-3000(-4300) m. Chongqing, S Gansu, Guangxi, Guizhou, Hebei, Henan, Liaoning, Shaanxi, Shanxi, Sichuan, Xizang [N India, Korea, Mongolia, Nepal, Russia (Siberia), Thailand, Vietnam].
别名：一把抓;还魂草;莪曲森得莫;九死还魂草;卷柏;老虎爪;神仙一把抓;石花;万年松;长生草;茧北;长生不死草;一反抓;石莲花;
科名：卷柏科 Selaginellaceae
属名：卷柏属 Selaginella
7.垫状卷柏（中国主要植物图说·蕨类植物门）图版23：1-8
Selaginella pulvinata (Hook. et Grev.) Maxim. in Mem. Acad. Imp. Sci. Petersb. 9: 335. 1859; Hand. -Mazz., Symb. Sin. 6: 5. 1929; Reed, Index Selaginellarum, inMem. Soc. Brot. 18: 187. 1966; Ic. Corm. Sin. 1: 111. 1972; Fl. Fujian. 1: 11. 1982; Fl. Xizang. 1: 24. 1983: B. D. Liu et al. in Bull. Bot. Res. (Harbin) 9 (3): 114, Pl. 5: 1-2. 1989; P. S. Wang in J. Arnold Arbor. 71: 265. 1990: R. D. Dixit, Selag. India 51, f. 16, Pl. 17. 1992; Fl. Jiangxi 1: 31, f. 16. 1993; Vasc. PI. Heng-duan Mts. 1: 9. 1993: P. S. Wang et X. Y. Wang, Pterid. Fl. Guizhou 639, pl. 155, f. 11-12. 2001. ——Lycopodium pulvinatum Hook. et Grev. in Hook. Bot. Misc. 2: 381. 1831. ——S. tamariscina var. pulvinata (Hook. et Grev.) Alston in Bull. FanMem. Inst. Biol. 5: 271. 1934, et in Proc. Nat. Inst. Sci. India 11: 221. 1945; S. H. Fu, Ill. Important Chinese Pl., Pterid. 8. 1957; Reed, Index Selaginellarum, in Mem.Soc. Brot. 18: 223. 1966; Fl. Henan 1: 5. 1981: R. D. Dixit, Cens. Indian Pterid. 16. 1984; Fl. Intramong. 1: 55, Pl. 2, f. 18. 1985: Fl. Liaoning. 1: 15. 1988; J. z. Wang et S. X. Li, Clav. Pl. Chin. Bor. -Ori. 2nd ed. 22. 1995. ——Lycopodioides pulv-inata (Hook. et Grev.) H. S. Kung, Fl. Sichuan. 6: 64, pl. 18, f. 1-3. 1988. ——S. involvens auct. non (Sw.) Spring: Baker, Handb. Fern-Allies 87, no. 204. 1887, p. p. : Warb. in Monsunia 1: 113. 1900, P. P. quoad pl. Peking; O. C. Schmidt in ActaHorti Gothob. 5: 52. 1930. ——S. tamariscina auct. non (P. Beauv.) Spring in Alstonin Lecomte, Fl. Gen. Indo-Chine 7 (2): 559. 1951; P. H. Ho, Ill. Fl. Vietnam 1: 27, cum fig. 1991.
土生或石生，旱生复苏植物，呈垫状，无匍匐根状茎或游走茎。根托只生于茎的基部，长2-4厘米，直径0.2-0.4毫米，根多分叉，密被毛，和茎及分枝密集形成树状主干，高数厘米。主茎自近基部羽状分枝，不呈“之”字形，禾秆色或棕色，主茎下部直径1毫米，不具沟槽，光滑，维管束1条； 侧枝4-7对，2-3回羽状分枝，小枝排列紧密，主茎上相邻分枝相距约1厘米，分枝无毛，背腹压扁，主茎在分枝部分中部连叶宽2.2-2.4毫米，末回分枝连叶宽1.2-1.6毫米。叶全部交互排列，二形，叶质厚，表面光滑，不具白边，主茎上的叶略大于分枝上的叶，相互重叠，绿色或棕色，斜升，边缘撕裂状。分枝上的腋叶对称，卵圆形到三角形，2.5毫米X 1.0毫米，边缘撕裂状并具睫毛。小枝上的叶斜卵形或三角形，2.8-3.1毫米XO.9-1.2毫米，覆瓦状排列，背部不呈龙骨状，先端具芒，基部平截（具簇毛），边缘撕裂状，并外卷。侧叶不对称，小枝上的叶距圆形，略斜升，2.9-3.2毫米X1.4-1.5毫米，先端具芒，边缘全缘，基部上侧扩大，加宽，覆盖小枝，基部上侧边缘不为全缘，呈撕裂状，基部下侧不呈耳状，边缘不为全缘，呈撕裂状，下侧边缘内卷。孢子叶穗紧密，四棱柱形，单生于小枝末端，10-20毫米X 1.5-2.0毫米；孢子叶一形，不具白边，边缘撕裂状，具睫毛；大孢子叶分布于孢子叶穗下部的下侧或中部的下侧或上部的下侧。大孢子黄白色或深褐色；小孢子浅黄色。
产山西、北京、重庆（城口、巫溪、秀山）、福建（德化、南靖）、甘肃（宕昌、文县）、广西（德宝、隆林）、贵州（册亨、从江、大方、关岭、贵阳、赫章、晴隆、黔西、水城、松桃、铜仁、威宁、西秀、兴义、镇宁）、河北（阜平、临城、山海关、蔚县、行唐、邢台、元氏、琢鹿）、河南（内乡）、江西（赣北）、辽宁（凌源）、陕西（眉县）、四川（安远、宝兴、北川、布拖、丹巴、道孚、德昌、都江堰、峨边、甘洛、高县、汉源、黑水、会理、剑阁、金川、金阳、九龙、康定、雷波、里塘、理县、芦山、泸定、马尔康、茂县、米易、冕宁、木里、南坪、宁南、平武、青川、石棉、汶川、西昌、喜德、乡里、小金、新龙、雅江、盐边、越西）、台湾、西藏（昌都、甘孜、拉萨、朗县、林周、芒康、米林、盐井）、云南（德钦、贡山、昆明、中甸）。常见于石灰岩上，海拔（100-) 1000-3000 (-4250)米。也分布到蒙古、俄罗斯西伯利亚、朝鲜半岛、日本、印度北部、越南、泰国等地。模式标本采自印度。
《Flora of China》 Vol. 2-3 (2013)
Selaginella pulvinata (Hooker & Greville) Maximowicz Mém. Acad. Imp. Sci. Saint Pétersbourg, sér. 7. 9: 335. 1859.
垫状卷柏 dian zhuang juan bai
Lycopodium pulvinatum Hooker & Greville, Hooker’s J. Bot. Kew Gard. Misc. 2: 381. 1831; Lycopodioides pulvinata (Hooker & Greville) H. S. Kung; Selaginella tamariscina (P. Beauvois) Spring var. pulvinata (Hooker & Greville) Alston.
Plants terrestrial or epilithic, xerophytic, “resurrectional”; stems forming rosette, seasonally green, stems and roots entangled forming treelike trunk, plants 2-10 cm, without creeping rhizomes. Rhizophores restricted to base of stem, 2-4 cm, rather thick or slender, much forked at end and forming thick massive rootstock. Main stems branched near and above base, pinnately branched, stramineous or brown, main stem ca. 1 mm in diam. at lower part; primary leafy branches 4-7 pairs, 2 or 3 times pinnately branched, branchlets dense, adjacent primary branches on main stem ca. 1 cm apart, leafy portion of main stem including leaves 2.2-2.4 mm wide at middle, ultimate branches 1.2-1.6 mm wide including leaves. Axillary leaves on main stems larger than those on branches; axillary leaves on branches symmetrical, ovate to triangular, ca. 2.5 × 1 mm, base exauriculate, margin lacerate-ciliolate. Dorsal leaves on branches imbricate, obliquely ovate or triangular, 2.8-3.1 × 0.9-1.2 mm, not carinate, base truncate (with a tuft of hairs), margin lacerate, outer margin revolute, apex aristate. Ventral leaves slightly ascending, asymmetrical, oblong, 2.9-3.2 × 1.4-1.5 mm, apex aristate, entire; basiscopic base not auriculate, margin lacerate, lower margin involute; acroscopic base enlarged, broader, overlapping stem and branches, margin lacerate. Strobili solitary, terminal, compact, tetragonal, 10-20 × 1.5-2 mm; sporophylls uniform, not white-margined, margin lacerate-ciliolate; megasporophylls in basal portion of strobili on lower side, in middle on lower side, or in upper portion on lower side; microspores pale yellow, megaspores white-yellow or dark brown.
Common in rock crevices on exposed rocky (limestone) ridges; (100-)1000-3000(-4300) m. Chongqing, S Gansu, Guangxi, Guizhou, Hebei, Henan, Liaoning, Shaanxi, Shanxi, Sichuan, Xizang [N India, Korea, Mongolia, Nepal, Russia (Siberia), Thailand, Vietnam].
