伏地卷柏Selaginella nipponica Franch. et Sav.
伏地卷柏Selaginella nipponica Franch. et Sav.
62.伏地卷柏（中国高等植物图鉴）图版65: 1-7
Selaginella nipponica Franch. et Sav., Enum. Pl. Jap. 2: 199, 615. 1879; Baker, Handh Fern-Allies 66. 1887; Komatsu et Matsumura, Ic. Pl. Koisikav. 1: Pl. 81. 1913; Alston in Bull. Fan Mem. Inst. Biol. 5: 272. 1934, p. p. ; S. H. Fu, Ill. Im-portant Chinese Pl., Pterid. 11, f. 15. 1957; Steward, Man. Vasc. Pl. Low. YangtzeVall. China 25. 1958; Makino, III. Fl. Jap. f. 17. 1961: Reed, Index Selaginellanam, in Mem. Soc. Brot. 18: 165. 1966; Ic. Corm. Sin. 1: 114, f. 228. 1972: Fl. Tsinling. 2: 21, Pl. 3, f. 1-6. 1974; M. Park, Ill. Encycl. Fauna et Fl. Kor. Vol. 16, Pterid. 116, 420. 1975; Fl. Jiangsu 1: 14, f. 7. 1977; Fl. Henan 1: 8, f. 11. 1981; T. C.Huang, Spore Fl. Taiwan 18, Pl. 9: 3 et pl. 15: 3. 1981; Fl. Fujian. 1: 12, f. 3. 1982; Fl. Anhui 1: 26, f. 11. 1986; B. D. Liu et -al. in Bull. Bot. Res. (Harbin) 9 (3): 114, Pl. 7: 2-4. 1989: P. S. Wang in J. Arnold Arbor. 71: 267. 1990; Y. L.Zhang et al., Spore Morph. Chin. Pterid. 46, f. 8c, Pl. 2: 39-42. 1990; Sa. Kurataet Nakaike, Ill. Pterid. Jap. 6: 178, cum photo. fig. et map. 1990: Nakaike, New Fl. Jap. Pterid. f. 44. 1992, et in J. Nat. Hist. Mus. Inst. Chiba 4 (2): 116. 1997: P. S.Wang, Pterid. Fanjing Mt. Nat. Res. 25. 1992; K. Iwats., Ferns Fern Allies Japan 53, Pl. 10, photo. 1-2. 1992, et K. Iwats., Fl. Jap. 1: 15. 1995; Fl. Jiangxi 1: 34, f. 21. 1993; Fl. Zhejiang 1: 17, f. 1-18. 1993: Vasc. Pl. Hengduan Mts. 1: 10. 1993; J. L. Tsai et W. C. Shieh in T. C. Huang, Fl. Taiwan ed. 2, 1: 53. 1994; P. S. Wang et X. Y. Wang, Pterid. Fl. Guizhou in T. C. Huang, 638. 2001. ——Lycopodioides nipponica Kuntze, Revis. Gen. Pl. 1: 827. 1891; H. S. Kung, Fl. Sichuan. 6: 62. 1988; Z. Y. Sun et al. in X. C. Zhang et K. H. Shing, Ching Mem.Vol. 342. 1999. ——S. savatieri Baker in J. Bot. (Hooker) 22: 87. 1884, et Handb. Fern-Allies 66. 1887; Baker in J. Bot. (Hooker) 26: 231. 1888; Warb. in Monsunia 1: 192. 1900: Diels in Engl. Bot. Jahrb. Syst. 29: 211. 1901; Hand. -Mazz., Symb. Sin. 6: 8. 1929; Ching in Sinensia 3 (12): 347. 1933: S. H. Fu, Ill. Important Chinese Pl., Pterid. 11. 1957, in syn.; De Vol et H. W. Chen in Taiwania 12: 82. 1966, pro syn. ; Reed, Index Selaginellarum, in Mem. Soc. Brot. 18: 203. 1966. ——Lycopodioides sa－vatieri (Baker) Kuntze, Revis. Gen. Pl. 1: 827. 1891. ——S. shensiensis Christ in Nuo-vo Giorn. Bot. Ital. n. s. 4: 102. 1897, p. p. : Reed, Index Selaginellarum, in Mem, Soc. Brot. 18: 209. 1966, p. p. ——S. hachijoensis Nakai in Bot. Mag. (Tokyo) 35: 149. 1921; Reed, Index Selaginellarum, in Mem. Soc. Brot. 18: 121. 1966; Nakaike, New Fl. Jap. Pterid. 44. 1992, pro syn. ——Urostachys ihwangensis Nessel, Fedde, Repert. Sp. Nov. Regni Veg. 39: 68, t. 192b. 1935, et in Sinensia 6: 664. 1935.
土生，匍匐，能育枝直立，高5-12厘米，无游走茎。根托沿匍匐茎和枝断续生长，自茎分叉处下方生出，长1-2.7厘米，纤细，直径0.1毫米，根少分叉，无毛。茎自近基部开始分枝，不呈“之”字形，无关节，禾秆色，茎下部直径0.2-0.4毫米，具沟槽，无毛，维管束1条；侧枝3-4对，不分叉或分叉或1回羽状分枝，分枝稀疏，茎上相邻分枝相距1-2厘米，叶状分枝和茎无毛，背腹压扁，茎在分枝部分中部连叶宽4.5-5.4毫米，末回分枝连叶宽2.8-4.2毫米。叶全部交互排列，二形，草质，表面光滑，边缘非全缘，不具白边。分枝上的腋叶对称或不对称，1.5-1.8毫米×0.8-1.0毫米，边缘有细齿。中叶多少对称，分枝上的中叶长圆状卵形或卵形或卵状披针形或椭圆形，1.6-2.0毫米×0.6一0.9毫米，紧接到覆瓦状（在先端部分）排列，背部不呈龙骨状，先端具尖头和急尖，基部钝，边缘不明显具细齿。侧叶不对称，侧枝上的侧叶宽卵形或卵状三角形，常反折，1.8-2.2毫米×1.0-1.6毫米，先端急尖；上侧基部扩大，加宽，覆盖小枝，上侧基部边缘具微齿。孢子叶穗疏松，通常背腹压扁，单生于小枝末端，或1-2 (3)次分叉，18-50毫米×2.0-4.6毫米；孢子叶二形或略二形，正置，和营养叶近似，排列一致，不具白边，边缘具细齿，背部不呈龙骨状，先端渐尖；大孢子叶分布于孢子叶穗下部的下侧。大孢子橘黄色；小孢子橘红色。
安徽（和县、霍山、青阳、休宁）、重庆（南川、石柱、秀山）、福建（崇安、泰宁、厦门）、甘肃（迭部、文县）、广东、广西（全县、融水）、贵州（安顺、岑巩、赤水、道真、德江、风岗、贵定、江口、仁怀、三穗、石阡、绥阳、天柱、务川、修文、沿河、玉屏、镇远、正安）、河南（大别山、灵宝、卢氏、栾川、洛宁、内乡、嵩县、桐柏山、西峡、淅川）、湖北（房县、兴山）、湖南（大庸、古丈、石门、宜章、永顺）、江苏（南京、苏州、镇江）、江西（安远、龙南、庐山、南昌、南丰、萍乡、新建、宜黄）、青海、陕西（长安、眉县、宁陕、平利）、山东、山西、四川（安县、成都、丹巴、道孚、德阳、都江堰、峨边、峨眉山、都江堰、广元、汉源、乐山、理县、茂县、绵阳、平武、三台、射洪、威远、雅安、雅江）、重庆（涪陵）、台湾（花莲、南投）、西藏（波密）、香港、云南（德钦、贡山、维西、中甸）、浙江（天台、雁荡山、舟山）。生于草地或岩石上，海拔80-1300米。也分布到日本。模式标本采自日本。
《Flora of China》 Vol. 2-3 (2013)
Selaginella nipponica Franchet & Savatier Enum. Pl. Jap. 2: 199, 615. 1879.
伏地卷柏 fu di juan bai
Lycopodioides nipponica (Franchet & Savatier) Kuntze; L. savatieri (Baker) Kuntze; Selaginella hachijoense Nakai; S. savatieri Baker; S. shensiensis Christ; Urostachys ihwangensis Nessel.
Plants terrestrial, seasonally green, creeping; fertile stems erect, 5-12 cm, without stolons, without elongate tuber at base of stem. Rhizophores restricted to sterile prostrate stems and branches at intervals throughout length of main stem, borne on ventral side in axils of branches. Main stems branched from near base upward, stramineous, 0.2-0.4 mm in diam. in lower part, angulate, sulcate; primary leafy branches 3 or 4 pairs, simple or forked or once pinnately branched, branchlets sparse, adjacent primary branches on main stem 1-2 cm apart, leafy branches and stems glabrous, dorsiventrally flattened, leafy portion of main stem including leaves 4.5-5.4 mm wide at middle, ultimate branches 2.8-4.2 mm wide including leaves. Axillary leaves on branches of sterile stems symmetrical or asymmetrical, ovate-lanceolate or elliptic, 1.5-1.8 × 0.8-1 mm, base exauriculate, margin denticulate. Dorsal leaves ± symmetrical, those on main stems not obviously larger than those on branches; dorsal leaves on branches contiguous or imbricate in apical portion, oblong-ovate, ovate, ovate-lanceolate, or elliptic, 1.6-2 × 0.6-0.9 mm, not carinate, base obtuse, margin not obviously denticulate, apex acuminate or acute. Ventral leaves asymmetrical, those on main stem not obviously larger than those on branches; ventral leaves on branches often deflexed, broadly ovate or ovate-triangular, 1.8-2.2 × 1-1.6 mm, apex acute; acroscopic base enlarged, broader, overlapping stem and branches, minutely denticulate. Fertile branches erect, 2.5-12 cm including strobilus. Strobili solitary or once or twice forked (or 3 times forked), terminal, lax, usually dorsiventrally complanate, 18-50 × 2-4.6 mm; sporophylls ± similar to sterile leaves in form and arrangement, dimorphic or slightly dimorphic, usually non-resupinate, not keeled, not white-margined, margin denticulate, apex acuminate; megasporophylls in basal portion on lower side of strobilus; microsporangia elliptic, relatively thick, with relatively large cells in central part; microspores orange-red, megaspores yellowish orange.
Grasslands, on rocks; 100-1300 m. Anhui, Chongqing, Fujian, S Gansu, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Qinghai, Shaanxi, Shandong, Shanxi, Sichuan, Taiwan, Yunnan, Zhejiang [Japan].
别名：接筋藤;小地柏;日本卷柏;地柏枝;异穗卷柏;石打穿;宽叶卷柏;补地云;铺地蜈蚣;
科名：卷柏科 Selaginellaceae
属名：卷柏属 Selaginella
62.伏地卷柏（中国高等植物图鉴）图版65: 1-7
Selaginella nipponica Franch. et Sav., Enum. Pl. Jap. 2: 199, 615. 1879; Baker, Handh Fern-Allies 66. 1887; Komatsu et Matsumura, Ic. Pl. Koisikav. 1: Pl. 81. 1913; Alston in Bull. Fan Mem. Inst. Biol. 5: 272. 1934, p. p. ; S. H. Fu, Ill. Im-portant Chinese Pl., Pterid. 11, f. 15. 1957; Steward, Man. Vasc. Pl. Low. YangtzeVall. China 25. 1958; Makino, III. Fl. Jap. f. 17. 1961: Reed, Index Selaginellanam, in Mem. Soc. Brot. 18: 165. 1966; Ic. Corm. Sin. 1: 114, f. 228. 1972: Fl. Tsinling. 2: 21, Pl. 3, f. 1-6. 1974; M. Park, Ill. Encycl. Fauna et Fl. Kor. Vol. 16, Pterid. 116, 420. 1975; Fl. Jiangsu 1: 14, f. 7. 1977; Fl. Henan 1: 8, f. 11. 1981; T. C.Huang, Spore Fl. Taiwan 18, Pl. 9: 3 et pl. 15: 3. 1981; Fl. Fujian. 1: 12, f. 3. 1982; Fl. Anhui 1: 26, f. 11. 1986; B. D. Liu et -al. in Bull. Bot. Res. (Harbin) 9 (3): 114, Pl. 7: 2-4. 1989: P. S. Wang in J. Arnold Arbor. 71: 267. 1990; Y. L.Zhang et al., Spore Morph. Chin. Pterid. 46, f. 8c, Pl. 2: 39-42. 1990; Sa. Kurataet Nakaike, Ill. Pterid. Jap. 6: 178, cum photo. fig. et map. 1990: Nakaike, New Fl. Jap. Pterid. f. 44. 1992, et in J. Nat. Hist. Mus. Inst. Chiba 4 (2): 116. 1997: P. S.Wang, Pterid. Fanjing Mt. Nat. Res. 25. 1992; K. Iwats., Ferns Fern Allies Japan 53, Pl. 10, photo. 1-2. 1992, et K. Iwats., Fl. Jap. 1: 15. 1995; Fl. Jiangxi 1: 34, f. 21. 1993; Fl. Zhejiang 1: 17, f. 1-18. 1993: Vasc. Pl. Hengduan Mts. 1: 10. 1993; J. L. Tsai et W. C. Shieh in T. C. Huang, Fl. Taiwan ed. 2, 1: 53. 1994; P. S. Wang et X. Y. Wang, Pterid. Fl. Guizhou in T. C. Huang, 638. 2001. ——Lycopodioides nipponica Kuntze, Revis. Gen. Pl. 1: 827. 1891; H. S. Kung, Fl. Sichuan. 6: 62. 1988; Z. Y. Sun et al. in X. C. Zhang et K. H. Shing, Ching Mem.Vol. 342. 1999. ——S. savatieri Baker in J. Bot. (Hooker) 22: 87. 1884, et Handb. Fern-Allies 66. 1887; Baker in J. Bot. (Hooker) 26: 231. 1888; Warb. in Monsunia 1: 192. 1900: Diels in Engl. Bot. Jahrb. Syst. 29: 211. 1901; Hand. -Mazz., Symb. Sin. 6: 8. 1929; Ching in Sinensia 3 (12): 347. 1933: S. H. Fu, Ill. Important Chinese Pl., Pterid. 11. 1957, in syn.; De Vol et H. W. Chen in Taiwania 12: 82. 1966, pro syn. ; Reed, Index Selaginellarum, in Mem. Soc. Brot. 18: 203. 1966. ——Lycopodioides sa－vatieri (Baker) Kuntze, Revis. Gen. Pl. 1: 827. 1891. ——S. shensiensis Christ in Nuo-vo Giorn. Bot. Ital. n. s. 4: 102. 1897, p. p. : Reed, Index Selaginellarum, in Mem, Soc. Brot. 18: 209. 1966, p. p. ——S. hachijoensis Nakai in Bot. Mag. (Tokyo) 35: 149. 1921; Reed, Index Selaginellarum, in Mem. Soc. Brot. 18: 121. 1966; Nakaike, New Fl. Jap. Pterid. 44. 1992, pro syn. ——Urostachys ihwangensis Nessel, Fedde, Repert. Sp. Nov. Regni Veg. 39: 68, t. 192b. 1935, et in Sinensia 6: 664. 1935.
土生，匍匐，能育枝直立，高5-12厘米，无游走茎。根托沿匍匐茎和枝断续生长，自茎分叉处下方生出，长1-2.7厘米，纤细，直径0.1毫米，根少分叉，无毛。茎自近基部开始分枝，不呈“之”字形，无关节，禾秆色，茎下部直径0.2-0.4毫米，具沟槽，无毛，维管束1条；侧枝3-4对，不分叉或分叉或1回羽状分枝，分枝稀疏，茎上相邻分枝相距1-2厘米，叶状分枝和茎无毛，背腹压扁，茎在分枝部分中部连叶宽4.5-5.4毫米，末回分枝连叶宽2.8-4.2毫米。叶全部交互排列，二形，草质，表面光滑，边缘非全缘，不具白边。分枝上的腋叶对称或不对称，1.5-1.8毫米×0.8-1.0毫米，边缘有细齿。中叶多少对称，分枝上的中叶长圆状卵形或卵形或卵状披针形或椭圆形，1.6-2.0毫米×0.6一0.9毫米，紧接到覆瓦状（在先端部分）排列，背部不呈龙骨状，先端具尖头和急尖，基部钝，边缘不明显具细齿。侧叶不对称，侧枝上的侧叶宽卵形或卵状三角形，常反折，1.8-2.2毫米×1.0-1.6毫米，先端急尖；上侧基部扩大，加宽，覆盖小枝，上侧基部边缘具微齿。孢子叶穗疏松，通常背腹压扁，单生于小枝末端，或1-2 (3)次分叉，18-50毫米×2.0-4.6毫米；孢子叶二形或略二形，正置，和营养叶近似，排列一致，不具白边，边缘具细齿，背部不呈龙骨状，先端渐尖；大孢子叶分布于孢子叶穗下部的下侧。大孢子橘黄色；小孢子橘红色。
安徽（和县、霍山、青阳、休宁）、重庆（南川、石柱、秀山）、福建（崇安、泰宁、厦门）、甘肃（迭部、文县）、广东、广西（全县、融水）、贵州（安顺、岑巩、赤水、道真、德江、风岗、贵定、江口、仁怀、三穗、石阡、绥阳、天柱、务川、修文、沿河、玉屏、镇远、正安）、河南（大别山、灵宝、卢氏、栾川、洛宁、内乡、嵩县、桐柏山、西峡、淅川）、湖北（房县、兴山）、湖南（大庸、古丈、石门、宜章、永顺）、江苏（南京、苏州、镇江）、江西（安远、龙南、庐山、南昌、南丰、萍乡、新建、宜黄）、青海、陕西（长安、眉县、宁陕、平利）、山东、山西、四川（安县、成都、丹巴、道孚、德阳、都江堰、峨边、峨眉山、都江堰、广元、汉源、乐山、理县、茂县、绵阳、平武、三台、射洪、威远、雅安、雅江）、重庆（涪陵）、台湾（花莲、南投）、西藏（波密）、香港、云南（德钦、贡山、维西、中甸）、浙江（天台、雁荡山、舟山）。生于草地或岩石上，海拔80-1300米。也分布到日本。模式标本采自日本。
《Flora of China》 Vol. 2-3 (2013)
Selaginella nipponica Franchet & Savatier Enum. Pl. Jap. 2: 199, 615. 1879.
伏地卷柏 fu di juan bai
Lycopodioides nipponica (Franchet & Savatier) Kuntze; L. savatieri (Baker) Kuntze; Selaginella hachijoense Nakai; S. savatieri Baker; S. shensiensis Christ; Urostachys ihwangensis Nessel.
Plants terrestrial, seasonally green, creeping; fertile stems erect, 5-12 cm, without stolons, without elongate tuber at base of stem. Rhizophores restricted to sterile prostrate stems and branches at intervals throughout length of main stem, borne on ventral side in axils of branches. Main stems branched from near base upward, stramineous, 0.2-0.4 mm in diam. in lower part, angulate, sulcate; primary leafy branches 3 or 4 pairs, simple or forked or once pinnately branched, branchlets sparse, adjacent primary branches on main stem 1-2 cm apart, leafy branches and stems glabrous, dorsiventrally flattened, leafy portion of main stem including leaves 4.5-5.4 mm wide at middle, ultimate branches 2.8-4.2 mm wide including leaves. Axillary leaves on branches of sterile stems symmetrical or asymmetrical, ovate-lanceolate or elliptic, 1.5-1.8 × 0.8-1 mm, base exauriculate, margin denticulate. Dorsal leaves ± symmetrical, those on main stems not obviously larger than those on branches; dorsal leaves on branches contiguous or imbricate in apical portion, oblong-ovate, ovate, ovate-lanceolate, or elliptic, 1.6-2 × 0.6-0.9 mm, not carinate, base obtuse, margin not obviously denticulate, apex acuminate or acute. Ventral leaves asymmetrical, those on main stem not obviously larger than those on branches; ventral leaves on branches often deflexed, broadly ovate or ovate-triangular, 1.8-2.2 × 1-1.6 mm, apex acute; acroscopic base enlarged, broader, overlapping stem and branches, minutely denticulate. Fertile branches erect, 2.5-12 cm including strobilus. Strobili solitary or once or twice forked (or 3 times forked), terminal, lax, usually dorsiventrally complanate, 18-50 × 2-4.6 mm; sporophylls ± similar to sterile leaves in form and arrangement, dimorphic or slightly dimorphic, usually non-resupinate, not keeled, not white-margined, margin denticulate, apex acuminate; megasporophylls in basal portion on lower side of strobilus; microsporangia elliptic, relatively thick, with relatively large cells in central part; microspores orange-red, megaspores yellowish orange.
Grasslands, on rocks; 100-1300 m. Anhui, Chongqing, Fujian, S Gansu, Guangdong, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Qinghai, Shaanxi, Shandong, Shanxi, Sichuan, Taiwan, Yunnan, Zhejiang [Japan].
