异穗卷柏Selaginella heterostachys Baker
异穗卷柏Selaginella heterostachys Baker
52.异穗卷柏（中国主要植物图说·蕨类植物门）图版57：1-9
Selaginella heterostachys Baker in J. Bot. (Hooker) 23: 177. 1885, et Handb. Fern-Allies: 110. 1887: Alston in Bull Fan Mem. Inst. 5: 290. 1934, et in Lecornte, F1. Gen. Indo-Chine 7 (2): 590. 1951; S. H. Fu, Ill. Important Chinese Pl., Pterid. 11. 1957; Steward, Man. Vasc. Pl. Low. Yangtze Vall. China 25. 1958; Tagawa, Col. Ill. Jap. Pterid. 255. 1959; De Vol et H. W. Chen in Taiwania 12: 89. 1966; Reed, In-dex Selaginellarum, in Mem. Soc. Brot. 18: 125. 1966: Ic. Corm. Sin. 1: 115. 1972; T. C. Huang, Spore Fl. Taiwan 17. 1981: Fl. Fujian. 1: 13. 1982: M. Minaki inPollen et Spores 26: 443, pl. 1: 4; Pl. 10: 1-3. 1984: Fl. Anhui 1: 27, f. 13. 1986; B. D. Liu et al. in Bull. Bot. Res. (Harbin) 9 (3): 114, Pl. 7: 5-7. 1989: P. S.Wang in J. Arnold Arbor. 71: 266. 1990; Y, L. Zhang et al., Spore Morph. Chin. Pterid. 47, Pl. 4: 11-13. 1990; Sa. Kurata et Nakaike, Ill. Pterid. Jap. 6: 140, cumphoto, fig. et map. 1990; P. H. Ho, Ill. Fl. Vietnam 1: 37, cum fig. 1991; P. S. Wang in Pterid. Fanjing Mt. Nat. Res. 24. 1992; Nakaike, New Fl. Jap. Pterid. f. 38. 1992; K. Iwats., Ferns Fern Allies Japan 54, Pl. 12, photo. 1-2. 1992, et Fl. Jap. 1: 16. 1995: Fl. Zhejiang 1: 16. 1993; Fl. Jiangxi 1: 38, f. 27. 1993; Vasc. Pl. Hengduan Mts. 1: 10. 1993; J. L. Tsai et W. C. Shieh in T. C. Huang, Fl. Taiwaned. 2, 1: 50. 1994; Nakaike in J. Nat. Hist. Mus. Inst. Chiba 4 (2): 113. 1997; P. S. Wang et X. Y. Wang, Pterid. Fl. Guizhou 634. 2001. ——Lycopodioides het-erostachya (Baker) Kuntze, Rev. Gen. Pl. 1: 826. 1891: H. S. Kung, Fl. Sichuan. 6: 77. 1988. ——S. recurvifolia Warb. in Monsunia 1: 125. 1900; Reed, Index Selaginel-larum, in Mem. Soc. Brot. 18: 190. 1966. ——S. praticola Hand. -Mazz. in Sitzb. A-cad. Wiss. Wien, Math. -Nat. 59: 50. 1922, et Symb. Sin. 6: 10, t. 2, f. 3. 1929; S. H. Fu, Ill. Important Chinese Pl., Pterid. 11. 1957, pro syn. : Reed, Index Selaginel-larum, in Mem. Soc. Brot. 18: 184. 1966. ——S. tarokensis Yamamoto in J. Soc. Trop. Agr. Taikohu 3: 237. 1931; Reed, Index Selaginellarum, in Mem. Soc. Brot. 18: 224. 1966. ——S. hezhangensis P. S. Wang et X. Y. Wang in Bull. Bot. Res. (Harbin) 16 (4): 418. 1996; P. S. Wang et X. Y. Wang, Pterid. Fl. Guizhou 634. 2001, syn. nov. ——S. nipponica auct. non Franch. et Sav. M. Park, Ill. Encycl. Fauna et Fl. Kor. vol. 16, Pterid. 116, 420, Pl. 50, f. 17. 1975, p. p. quoad f.
土生或石生，直立或匍匐，直立能育茎高10-20厘米，具匍匐茎。根托沿匍匐茎断续孢生，但只生直立茎下部，自茎分叉处下方生出，长0.5-3.5厘米，纤细，直径0.1-0.3毫米，根少分叉，被毛。茎羽状分枝，不呈“之”字形，无关节，禾秆色，下部直径0.4-1.2毫米，茎圆柱状，具沟槽，无毛，维管束1条，直立能育茎自下部开始分枝，侧枝3-5对，1-2回羽状分枝，小枝稀疏，规则，茎上相邻分枝相距1.5-6厘米，分枝无毛，背腹压扁，茎在分枝部分中部连叶宽3-6毫米，末回分枝连叶宽2.4-5.6毫米。叶全部交互排列，二形，草质，表面光滑，无虹彩，边缘不为全缘，不具白边。茎上的腋叶较分枝上的大，卵圆形，近心形，分枝上的腋叶对称，卵形或长圆形，1.4-2.6毫米× 0.4-1.2毫米，边缘有细齿。中叶不对称，分枝上的中叶卵形或卵状披针形，1.0-1.6毫米×0.4-0.8毫米，相互排列不是非常近，背部不呈龙骨状，先端外展或与轴平行，先端具尖头或短芒，基部楔形，边缘具微齿。侧叶不对称，主茎上的明显大于侧枝上的，侧枝上的侧叶长圆状卵圆形，外展或下折，排列疏或密，1.8-2.7毫米×0.7-1.8毫米，先端急尖。边缘有细齿，上侧基部扩大，加宽，覆盖小枝，上侧基部边缘有细齿，下侧基部圆形，具细齿。孢子叶穗紧密，背腹压扁，单生于小枝末端，5-25毫米×1.5-3.5毫米；孢子叶明显二形，倒置，上侧的孢子叶卵状披针形或长圆状镰形，边缘具缘毛或具细齿，背部不呈龙骨状，先端具尖头到芒，上侧的孢子叶具孢子叶翼，孢子叶翼达叶尖，边缘具短睫毛或细齿，下侧的孢子叶卵状披针形，边缘具缘毛，先端具长尖头，龙骨状，脊上具睫毛；大孢子叶分布于孢子叶穗上下两侧的基部，或大、小孢子叶相间排列。大孢子橘黄色；小孢子橘黄色。
安徽（祁门、欺县、休宁）、重庆（南川、万县）、福建（崇安、德化、福州、将乐、南平、上杭、厦门、武夷）、甘肃（文县）、广东（深圳）、广西（罗城、天峨）、贵州（安顺、毕节、册亨、都匀、德江、贵定、贵阳、关岭、江口、荔波、龙里、雷山、三都、惠水、麻江、普定、绥阳、桐梓、务川、息峰、西秀、遵义、镇远、紫云、镇宁）、海南（昌江、陵水、琼中）、河南（内乡）、香港、湖南（古丈、石门、韶山、湘潭）、江西（定南、九江、龙南、庐山、铅山、资溪）、四川（都江堰、峨眉山、都江堰、汉源、泸定、简阳、剑阁、攀枝花、荥经、仁寿、天全）、台湾（高雄、花莲、屏东、南投、台东）、云南（贡山、安宁、双柏、马关、洱源）、浙江（昌化、杭州）。生于林下岩石上，海拔130-1300（-1900）米。模式标本采自香港。
《Flora of China》 Vol. 2-3 (2013)
Selaginella heterostachys Baker J. Bot. (Hooker). 23: 177. 1885.
异穗卷柏 yi sui juan bai
Lycopodioides heterostachya (Baker) Kuntze; Selaginella hezhangensis P. S. Wang & X. Y. Wang; S. praticola Handel-Mazzetti; S. recurvifolia Warburg; S. tarokoensis Yamamoto.
Plants terrestrial or epilithic, evergreen, erect or creeping, 10-20 cm, fertile stems erect. Rhizophores restricted to lower part of erect fertile branches or at intervals throughout length of creeping stem and branches, borne on ventral side in axils of branches. Main stem pinnately branched, stramineous, 0.4-1.2 mm in diam. in lower part, terete, sulcate, erect fertile stems branched from lower part upward; primary leafy branches 3-5 pairs, once or twice pinnately branched, secondary branches once or twice forked, tertiary branches forked or simple, branchlets sparse and regular, adjacent primary branches on main stem 1.5-6 cm apart, leafy portion of main stem including leaves 3-6 mm wide at middle, ultimate branches 2.4-5.6 mm wide including leaves. Axillary leaves on main stems larger than those on branches, ovate, base not peltate, subcordate; axillary leaves on branches symmetrical, ovate or oblong, 1.4-2.6 × 0.4-1.2 mm, base exauriculate, margin denticulate. Dorsal leaves asymmetrical, those on branches not approximate, ovate or ovate-lanceolate, 1-1.6 × 0.4-0.8 mm, not carinate, base cuneate or obliquely attenuate, not peltate, margin minutely denticulate, apex acuminate or shortly aristate. Ventral leaves asymmetrical, those on main stem obviously larger than those on branches; ventral leaves on branches distant or approximate, spreading or deflexed, oblong-ovate, 1.8-2.7 × 0.7-1.8 mm, margin denticulate, apex acute; basiscopic base rounded; acroscopic base enlarged, broader, overlapping stem and branches. Strobili solitary, terminal, compact, dorsiventrally complanate, 5-25 × 1.5-3.5 mm; sporophylls strongly dimorphic, resupinate; dorsal sporophylls ovate-lanceolate or oblong-lanceolate, not keeled, margin ciliolate or denticulate, apex acuminate or aristate, with sporophyll-pteryx complete and ciliolate or denticulate; ventral sporophylls ovate-lanceolate, carinate (ridge also ciliolate), margin ciliolate, apex long acuminate; megasporophylls in basal portion on both sides or lower side of strobilus, ± randomly distributed on both sides, or throughout lower side, or megasporophylls and microsporophylls at intervals; microsporangia transversely elliptic, rather thin, cells uniform; microspores yellowish orange, megaspores yellowish orange.
● On rocks in forests, wet habitats including on moss-covered or wet rocks and walls, occasionally on tree trunks close to streams; 100-1300(-1900) m. Anhui, Chongqing, Fujian, S Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangxi, Sichuan, Taiwan, Yunnan, Zhejiang.
别名：姬卷柏;火烫药;
科名：卷柏科 Selaginellaceae
属名：卷柏属 Selaginella
52.异穗卷柏（中国主要植物图说·蕨类植物门）图版57：1-9
Selaginella heterostachys Baker in J. Bot. (Hooker) 23: 177. 1885, et Handb. Fern-Allies: 110. 1887: Alston in Bull Fan Mem. Inst. 5: 290. 1934, et in Lecornte, F1. Gen. Indo-Chine 7 (2): 590. 1951; S. H. Fu, Ill. Important Chinese Pl., Pterid. 11. 1957; Steward, Man. Vasc. Pl. Low. Yangtze Vall. China 25. 1958; Tagawa, Col. Ill. Jap. Pterid. 255. 1959; De Vol et H. W. Chen in Taiwania 12: 89. 1966; Reed, In-dex Selaginellarum, in Mem. Soc. Brot. 18: 125. 1966: Ic. Corm. Sin. 1: 115. 1972; T. C. Huang, Spore Fl. Taiwan 17. 1981: Fl. Fujian. 1: 13. 1982: M. Minaki inPollen et Spores 26: 443, pl. 1: 4; Pl. 10: 1-3. 1984: Fl. Anhui 1: 27, f. 13. 1986; B. D. Liu et al. in Bull. Bot. Res. (Harbin) 9 (3): 114, Pl. 7: 5-7. 1989: P. S.Wang in J. Arnold Arbor. 71: 266. 1990; Y, L. Zhang et al., Spore Morph. Chin. Pterid. 47, Pl. 4: 11-13. 1990; Sa. Kurata et Nakaike, Ill. Pterid. Jap. 6: 140, cumphoto, fig. et map. 1990; P. H. Ho, Ill. Fl. Vietnam 1: 37, cum fig. 1991; P. S. Wang in Pterid. Fanjing Mt. Nat. Res. 24. 1992; Nakaike, New Fl. Jap. Pterid. f. 38. 1992; K. Iwats., Ferns Fern Allies Japan 54, Pl. 12, photo. 1-2. 1992, et Fl. Jap. 1: 16. 1995: Fl. Zhejiang 1: 16. 1993; Fl. Jiangxi 1: 38, f. 27. 1993; Vasc. Pl. Hengduan Mts. 1: 10. 1993; J. L. Tsai et W. C. Shieh in T. C. Huang, Fl. Taiwaned. 2, 1: 50. 1994; Nakaike in J. Nat. Hist. Mus. Inst. Chiba 4 (2): 113. 1997; P. S. Wang et X. Y. Wang, Pterid. Fl. Guizhou 634. 2001. ——Lycopodioides het-erostachya (Baker) Kuntze, Rev. Gen. Pl. 1: 826. 1891: H. S. Kung, Fl. Sichuan. 6: 77. 1988. ——S. recurvifolia Warb. in Monsunia 1: 125. 1900; Reed, Index Selaginel-larum, in Mem. Soc. Brot. 18: 190. 1966. ——S. praticola Hand. -Mazz. in Sitzb. A-cad. Wiss. Wien, Math. -Nat. 59: 50. 1922, et Symb. Sin. 6: 10, t. 2, f. 3. 1929; S. H. Fu, Ill. Important Chinese Pl., Pterid. 11. 1957, pro syn. : Reed, Index Selaginel-larum, in Mem. Soc. Brot. 18: 184. 1966. ——S. tarokensis Yamamoto in J. Soc. Trop. Agr. Taikohu 3: 237. 1931; Reed, Index Selaginellarum, in Mem. Soc. Brot. 18: 224. 1966. ——S. hezhangensis P. S. Wang et X. Y. Wang in Bull. Bot. Res. (Harbin) 16 (4): 418. 1996; P. S. Wang et X. Y. Wang, Pterid. Fl. Guizhou 634. 2001, syn. nov. ——S. nipponica auct. non Franch. et Sav. M. Park, Ill. Encycl. Fauna et Fl. Kor. vol. 16, Pterid. 116, 420, Pl. 50, f. 17. 1975, p. p. quoad f.
土生或石生，直立或匍匐，直立能育茎高10-20厘米，具匍匐茎。根托沿匍匐茎断续孢生，但只生直立茎下部，自茎分叉处下方生出，长0.5-3.5厘米，纤细，直径0.1-0.3毫米，根少分叉，被毛。茎羽状分枝，不呈“之”字形，无关节，禾秆色，下部直径0.4-1.2毫米，茎圆柱状，具沟槽，无毛，维管束1条，直立能育茎自下部开始分枝，侧枝3-5对，1-2回羽状分枝，小枝稀疏，规则，茎上相邻分枝相距1.5-6厘米，分枝无毛，背腹压扁，茎在分枝部分中部连叶宽3-6毫米，末回分枝连叶宽2.4-5.6毫米。叶全部交互排列，二形，草质，表面光滑，无虹彩，边缘不为全缘，不具白边。茎上的腋叶较分枝上的大，卵圆形，近心形，分枝上的腋叶对称，卵形或长圆形，1.4-2.6毫米× 0.4-1.2毫米，边缘有细齿。中叶不对称，分枝上的中叶卵形或卵状披针形，1.0-1.6毫米×0.4-0.8毫米，相互排列不是非常近，背部不呈龙骨状，先端外展或与轴平行，先端具尖头或短芒，基部楔形，边缘具微齿。侧叶不对称，主茎上的明显大于侧枝上的，侧枝上的侧叶长圆状卵圆形，外展或下折，排列疏或密，1.8-2.7毫米×0.7-1.8毫米，先端急尖。边缘有细齿，上侧基部扩大，加宽，覆盖小枝，上侧基部边缘有细齿，下侧基部圆形，具细齿。孢子叶穗紧密，背腹压扁，单生于小枝末端，5-25毫米×1.5-3.5毫米；孢子叶明显二形，倒置，上侧的孢子叶卵状披针形或长圆状镰形，边缘具缘毛或具细齿，背部不呈龙骨状，先端具尖头到芒，上侧的孢子叶具孢子叶翼，孢子叶翼达叶尖，边缘具短睫毛或细齿，下侧的孢子叶卵状披针形，边缘具缘毛，先端具长尖头，龙骨状，脊上具睫毛；大孢子叶分布于孢子叶穗上下两侧的基部，或大、小孢子叶相间排列。大孢子橘黄色；小孢子橘黄色。
安徽（祁门、欺县、休宁）、重庆（南川、万县）、福建（崇安、德化、福州、将乐、南平、上杭、厦门、武夷）、甘肃（文县）、广东（深圳）、广西（罗城、天峨）、贵州（安顺、毕节、册亨、都匀、德江、贵定、贵阳、关岭、江口、荔波、龙里、雷山、三都、惠水、麻江、普定、绥阳、桐梓、务川、息峰、西秀、遵义、镇远、紫云、镇宁）、海南（昌江、陵水、琼中）、河南（内乡）、香港、湖南（古丈、石门、韶山、湘潭）、江西（定南、九江、龙南、庐山、铅山、资溪）、四川（都江堰、峨眉山、都江堰、汉源、泸定、简阳、剑阁、攀枝花、荥经、仁寿、天全）、台湾（高雄、花莲、屏东、南投、台东）、云南（贡山、安宁、双柏、马关、洱源）、浙江（昌化、杭州）。生于林下岩石上，海拔130-1300（-1900）米。模式标本采自香港。
《Flora of China》 Vol. 2-3 (2013)
Selaginella heterostachys Baker J. Bot. (Hooker). 23: 177. 1885.
异穗卷柏 yi sui juan bai
Lycopodioides heterostachya (Baker) Kuntze; Selaginella hezhangensis P. S. Wang & X. Y. Wang; S. praticola Handel-Mazzetti; S. recurvifolia Warburg; S. tarokoensis Yamamoto.
Plants terrestrial or epilithic, evergreen, erect or creeping, 10-20 cm, fertile stems erect. Rhizophores restricted to lower part of erect fertile branches or at intervals throughout length of creeping stem and branches, borne on ventral side in axils of branches. Main stem pinnately branched, stramineous, 0.4-1.2 mm in diam. in lower part, terete, sulcate, erect fertile stems branched from lower part upward; primary leafy branches 3-5 pairs, once or twice pinnately branched, secondary branches once or twice forked, tertiary branches forked or simple, branchlets sparse and regular, adjacent primary branches on main stem 1.5-6 cm apart, leafy portion of main stem including leaves 3-6 mm wide at middle, ultimate branches 2.4-5.6 mm wide including leaves. Axillary leaves on main stems larger than those on branches, ovate, base not peltate, subcordate; axillary leaves on branches symmetrical, ovate or oblong, 1.4-2.6 × 0.4-1.2 mm, base exauriculate, margin denticulate. Dorsal leaves asymmetrical, those on branches not approximate, ovate or ovate-lanceolate, 1-1.6 × 0.4-0.8 mm, not carinate, base cuneate or obliquely attenuate, not peltate, margin minutely denticulate, apex acuminate or shortly aristate. Ventral leaves asymmetrical, those on main stem obviously larger than those on branches; ventral leaves on branches distant or approximate, spreading or deflexed, oblong-ovate, 1.8-2.7 × 0.7-1.8 mm, margin denticulate, apex acute; basiscopic base rounded; acroscopic base enlarged, broader, overlapping stem and branches. Strobili solitary, terminal, compact, dorsiventrally complanate, 5-25 × 1.5-3.5 mm; sporophylls strongly dimorphic, resupinate; dorsal sporophylls ovate-lanceolate or oblong-lanceolate, not keeled, margin ciliolate or denticulate, apex acuminate or aristate, with sporophyll-pteryx complete and ciliolate or denticulate; ventral sporophylls ovate-lanceolate, carinate (ridge also ciliolate), margin ciliolate, apex long acuminate; megasporophylls in basal portion on both sides or lower side of strobilus, ± randomly distributed on both sides, or throughout lower side, or megasporophylls and microsporophylls at intervals; microsporangia transversely elliptic, rather thin, cells uniform; microspores yellowish orange, megaspores yellowish orange.
● On rocks in forests, wet habitats including on moss-covered or wet rocks and walls, occasionally on tree trunks close to streams; 100-1300(-1900) m. Anhui, Chongqing, Fujian, S Gansu, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangxi, Sichuan, Taiwan, Yunnan, Zhejiang.
