假糙苏Paraphlomis javanica (Bl.) Prain
假糙苏Paraphlomis javanica (Bl.) Prain
1．假糙苏
Paraphlomis javanica (Bl.) Prain in Ann. Bot. Gard. Calc. 9: 59. 1901, in obs.; Back. & Bakh. f. Fl. Java 2: 619. 1965; H. Keng in Gard. Bull. Singapore 24: 136, f. 26, a-g. 1969——Leonurus javanicus Bl. Bijdr. 828. 1826; Benth. Labiat. Gen. et Sp. 522. 1834——Phlomis javanica (Bl. ) Prain in Ann. Bot. Gard. Calc. 3: 231. 1891; 9: 59. 1901——Phlomis rugosa Benth. in Wall. Cat. n. 2067. 1829, nom. nud., in Wall. Pl. Asiat, Rar. 1: 63. 1830, Labiat. Gen. et Sp. 634. 1836, et in Walp. Rep. 3: 889. 1845; Hook. f. Fl. Brit. Ind. 4: 693. 1885; Prain in Ann. Bot. Gard. Calc. 3: 231, 1891, in nota; Stapf in Trans. Linn. Soc. London 4 (2): 216. 1894; Dunn in Notes Bot. Gard. Edinburgh 8: 168-9. 1913, 6: 186. 1915, quoad specim. Yunnan. ——Paraphlomis rugosa (Benth. ) Prain in Ann. Bot. Gard. Calc. 9: 60, pl. 74. 1901, in nota; in Journ. Asiat. Soc. Bengal 74: 721. 1907; Merr. Enum. Philip. Fl. Pl. 3: 412. 1922; Ridley, Fl. Malay Penins. 2: 651. 1923; Merr. in Lingnan Agr. Rev. 2: 138. 1925; Kudo in Mem. Fac. Sci. Agr. Taihoku Univ. 2: 209. 1929, p. p.; Sasaki, Cat. Governm. Herb. 443. 1930; Masam. Short. Fl. Formosa 185. 1936; Doan in Lecte. Gen. Fl. Indo-Chine 4: 1015. 1936; Hand. -Mazz. Symb. Sin. 7: 922. 1936, et in Beih. Bot. Centralbl. 56B: 458. 1939; Y. Z. Sun in Act. Phytotax. Sin. 4 (1): 50. 1955, p. p. min.; C. Y. Wu in ibid. 8 (1): 37. 1959; H. W. Li in ibid. 10 (1): 60. 1965 Lamium longipetiolata Hayata, Icon. Pl. Formos. 8: 92. 1919.
形态特征：草本，从纤细须根上升。茎单生，通常高约50厘米，有时高达1．5米，钝四棱形，具槽，被倒向平伏毛，常曲折，向基部无叶，上部具叶。叶椭圆形、椭圆状卵形或长圆状卵形，通常长7—15厘米，宽3—8．5厘米，间或有时长达30厘米，宽达14厘米，先端锐尖或渐尖，基部圆形或近楔形，边缘有具小突尖的圆齿状锯齿，有时齿多少不明显，膜质或纸质，上面绿色，多少被小刚毛，下面淡绿色，沿脉上密生余部疏生平伏毛，侧脉5—6对，在上面不明显，下面稍隆起；叶柄纤弱，长达8厘米，扁平，上面略具槽，下面圆形，被平伏毛。轮伞花序多花，轮廓为圆球形，连花冠径约3厘米，其下承以少数小苞片；小苞片钻形，长约6毫米，不超过萼筒，被小硬毛；花梗无。花萼花时明显管状，口部骤然开张，果时膨大，革质，不连齿长约7毫米，幼时密被小硬毛，多少呈绿色，果时毛被近于脱落，常变红色，脉不明显，齿5，近相等，钻形或三角状钻形，长3—4毫米。花冠通常黄或淡黄，亦有近于白色的，长约1．7厘米，外面在冠筒上部及冠檐上均多少被有小硬毛，内面在冠筒的中部稍上方具柔毛毛环，冠檐二唇形，上唇长圆形，全缘，直伸，下唇3裂，中裂片较大。雄蕊4，前对较长，均上升至上唇片之下或略超出，花丝丝状，微被柔毛，花药椭圆形，二室，室略叉开。花柱丝状，略超出雄蕊，先端近相等2浅裂。子房紫黑色，顶端截平，中央稍凹陷，无毛。小坚果倒卵珠状三棱形，顶端钝圆，黑色，无毛。花期6—8月，果期8—12月。
产地分布：产云南东南部、南部及西南部，广西西南部，广东海南，台湾；生于热带林荫下，海拔320—1350(—2500)米。印度，巴基斯坦，缅甸，泰国，老挝，越南经马来西亚至印度尼西亚及菲律宾也有。模式标本采自印度尼西亚的爪哇。
参考文献：Paraphlomisjavaniea(Bl)PraininAnn．Bot．Gard，Calc．9：59．1901，inobs．；Back．&Bakh．f．Fl.Java2：619．1965；H．KenginGard．Bull．Singa-pore24：136，f.26，a—g.1969——LeonurusjavabucysBl．Bijdr．828．1826;Benth．Labial．Gen．etSp．522．1834——Phlomisjavanica(Bl．)PraininAnn．Bot．Garti．Calc．3：231．1891；9：59．1901——PhlomisrugosaBenth．inWall．Cat．n．2067．1829，nom．nud．inWall．Pl．Asiat．Rar．1：63．1830，Labiat．Gen．etSp.634．1836，etinWalp．Rep.3：889．1845；Hook．f．FlBrit．Ind．4：693．1885;PraininAnn．Bot．Gard．Calc．3：231，1891，innota；StapfinTrans．Linn．Soc．London4(2)：216．1894；DunninNotesBot．Card．Edinburgh8：168——9．1913，6：186．1915，quoadspecim．Yunnan．——Paraphlomisrugosa(Benth．)PraininAnn．Bot．Gard．Calc．9：60，Pl．74．1901，innota；inJourn．Asiat．Soc．Bengal74：721．1907；Merr．Fmum．Philip．FlPl3：412．1922；Ridley，FlMalayPenins．2：651．1923；Merr．inLingnanAgr．Rev．2：138．1925;KudoinMem．Fac．Sci．Agr．TaihokuUniv．2：209．1929，P．P.；Sasaki，Cat．Governm．Herb．443．1930；Masam．Short．Fl．Formosa185．1936；DoaninLecte．Gen．FlIndo-Chine4：1015．1936；Hand．-Mazz．Symb．Sin．7：922．1936，etinBeih．Bot．Centralbl．56B：458．1939；Y．Z．SuninAct．Phytotax．Sin．4(1)：50．1955，P．P．min．；c.Y．Wuinibid．8(1)：37．1959；H．W．1iinibid．10(1)：60．1965——LamiumlongipetiolataHayata，Icon．Pl．Formos．8：92．1919．
本系概述：Ser．JavanicaeC．Y．WuetH．WLiinAct．Phytotax．Sin．10(1)：60．1965．萼齿明显，宽至狭三角形，先端钻形，刚毛状渐尖或具短尖头。
《Flora of China》 Vol. 17 (1994)
Paraphlomis javanica (Blume) Prain
假糙苏 jia cao su
Herbs erect. Stem solitary, 0.5-1.5 m, retrorse strigose, flexuous, leafless toward base. Petiole slender, to 8 cm; leaf blade elliptic to oblong-ovate, 3-15(-30) × 1.5-8.5(-14) cm, adaxially ± minutely hispid, abaxially strigose, base rounded to subtruncate, margin inconspicuously serrate to crenate-serrate, apex acute to acuminate. Verticillasters many flowered, globose, ca. 3 cm in diam., surrounded by few bracteoles at base; bracteoles subulate, ca. 6 mm, shorter than calyx, hirtellous. Flowers sessile. Calyx tubular in flower, abruptly spreading at mouth, enlarged and red in fruit, leathery, densely hirtellous or minutely hispid, glabrescent, inconspicuously veined, tube ca. 7 mm; teeth subequal, needlelike to triangular, 2-4 mm. Corolla yellow or yellowish, less often whitish, ca. 1.7 cm, ± hirtellous on apical parts of tube and limb outside, pubescent annulate inside, upper lip oblong, entire. Nutlets black, triquetrous, obovoid, apex rounded, glabrous. Fl. Jun-Aug, fr. Aug-Dec.
Evergreen and mixed forests; 300-2400 m. Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangxi, Sichuan, Taiwan, Yunnan [India, Indonesia, Laos, Malaysia, Myanmar, Pakistan, Philippines, Thailand, Vietnam].
科名：唇形科 Labiatae
属名：假糙苏属 Paraphlomis
1．假糙苏
Paraphlomis javanica (Bl.) Prain in Ann. Bot. Gard. Calc. 9: 59. 1901, in obs.; Back. & Bakh. f. Fl. Java 2: 619. 1965; H. Keng in Gard. Bull. Singapore 24: 136, f. 26, a-g. 1969——Leonurus javanicus Bl. Bijdr. 828. 1826; Benth. Labiat. Gen. et Sp. 522. 1834——Phlomis javanica (Bl. ) Prain in Ann. Bot. Gard. Calc. 3: 231. 1891; 9: 59. 1901——Phlomis rugosa Benth. in Wall. Cat. n. 2067. 1829, nom. nud., in Wall. Pl. Asiat, Rar. 1: 63. 1830, Labiat. Gen. et Sp. 634. 1836, et in Walp. Rep. 3: 889. 1845; Hook. f. Fl. Brit. Ind. 4: 693. 1885; Prain in Ann. Bot. Gard. Calc. 3: 231, 1891, in nota; Stapf in Trans. Linn. Soc. London 4 (2): 216. 1894; Dunn in Notes Bot. Gard. Edinburgh 8: 168-9. 1913, 6: 186. 1915, quoad specim. Yunnan. ——Paraphlomis rugosa (Benth. ) Prain in Ann. Bot. Gard. Calc. 9: 60, pl. 74. 1901, in nota; in Journ. Asiat. Soc. Bengal 74: 721. 1907; Merr. Enum. Philip. Fl. Pl. 3: 412. 1922; Ridley, Fl. Malay Penins. 2: 651. 1923; Merr. in Lingnan Agr. Rev. 2: 138. 1925; Kudo in Mem. Fac. Sci. Agr. Taihoku Univ. 2: 209. 1929, p. p.; Sasaki, Cat. Governm. Herb. 443. 1930; Masam. Short. Fl. Formosa 185. 1936; Doan in Lecte. Gen. Fl. Indo-Chine 4: 1015. 1936; Hand. -Mazz. Symb. Sin. 7: 922. 1936, et in Beih. Bot. Centralbl. 56B: 458. 1939; Y. Z. Sun in Act. Phytotax. Sin. 4 (1): 50. 1955, p. p. min.; C. Y. Wu in ibid. 8 (1): 37. 1959; H. W. Li in ibid. 10 (1): 60. 1965 Lamium longipetiolata Hayata, Icon. Pl. Formos. 8: 92. 1919.
形态特征：草本，从纤细须根上升。茎单生，通常高约50厘米，有时高达1．5米，钝四棱形，具槽，被倒向平伏毛，常曲折，向基部无叶，上部具叶。叶椭圆形、椭圆状卵形或长圆状卵形，通常长7—15厘米，宽3—8．5厘米，间或有时长达30厘米，宽达14厘米，先端锐尖或渐尖，基部圆形或近楔形，边缘有具小突尖的圆齿状锯齿，有时齿多少不明显，膜质或纸质，上面绿色，多少被小刚毛，下面淡绿色，沿脉上密生余部疏生平伏毛，侧脉5—6对，在上面不明显，下面稍隆起；叶柄纤弱，长达8厘米，扁平，上面略具槽，下面圆形，被平伏毛。轮伞花序多花，轮廓为圆球形，连花冠径约3厘米，其下承以少数小苞片；小苞片钻形，长约6毫米，不超过萼筒，被小硬毛；花梗无。花萼花时明显管状，口部骤然开张，果时膨大，革质，不连齿长约7毫米，幼时密被小硬毛，多少呈绿色，果时毛被近于脱落，常变红色，脉不明显，齿5，近相等，钻形或三角状钻形，长3—4毫米。花冠通常黄或淡黄，亦有近于白色的，长约1．7厘米，外面在冠筒上部及冠檐上均多少被有小硬毛，内面在冠筒的中部稍上方具柔毛毛环，冠檐二唇形，上唇长圆形，全缘，直伸，下唇3裂，中裂片较大。雄蕊4，前对较长，均上升至上唇片之下或略超出，花丝丝状，微被柔毛，花药椭圆形，二室，室略叉开。花柱丝状，略超出雄蕊，先端近相等2浅裂。子房紫黑色，顶端截平，中央稍凹陷，无毛。小坚果倒卵珠状三棱形，顶端钝圆，黑色，无毛。花期6—8月，果期8—12月。
产地分布：产云南东南部、南部及西南部，广西西南部，广东海南，台湾；生于热带林荫下，海拔320—1350(—2500)米。印度，巴基斯坦，缅甸，泰国，老挝，越南经马来西亚至印度尼西亚及菲律宾也有。模式标本采自印度尼西亚的爪哇。
参考文献：Paraphlomisjavaniea(Bl)PraininAnn．Bot．Gard，Calc．9：59．1901，inobs．；Back．&Bakh．f．Fl.Java2：619．1965；H．KenginGard．Bull．Singa-pore24：136，f.26，a—g.1969——LeonurusjavabucysBl．Bijdr．828．1826;Benth．Labial．Gen．etSp．522．1834——Phlomisjavanica(Bl．)PraininAnn．Bot．Garti．Calc．3：231．1891；9：59．1901——PhlomisrugosaBenth．inWall．Cat．n．2067．1829，nom．nud．inWall．Pl．Asiat．Rar．1：63．1830，Labiat．Gen．etSp.634．1836，etinWalp．Rep.3：889．1845；Hook．f．FlBrit．Ind．4：693．1885;PraininAnn．Bot．Gard．Calc．3：231，1891，innota；StapfinTrans．Linn．Soc．London4(2)：216．1894；DunninNotesBot．Card．Edinburgh8：168——9．1913，6：186．1915，quoadspecim．Yunnan．——Paraphlomisrugosa(Benth．)PraininAnn．Bot．Gard．Calc．9：60，Pl．74．1901，innota；inJourn．Asiat．Soc．Bengal74：721．1907；Merr．Fmum．Philip．FlPl3：412．1922；Ridley，FlMalayPenins．2：651．1923；Merr．inLingnanAgr．Rev．2：138．1925;KudoinMem．Fac．Sci．Agr．TaihokuUniv．2：209．1929，P．P.；Sasaki，Cat．Governm．Herb．443．1930；Masam．Short．Fl．Formosa185．1936；DoaninLecte．Gen．FlIndo-Chine4：1015．1936；Hand．-Mazz．Symb．Sin．7：922．1936，etinBeih．Bot．Centralbl．56B：458．1939；Y．Z．SuninAct．Phytotax．Sin．4(1)：50．1955，P．P．min．；c.Y．Wuinibid．8(1)：37．1959；H．W．1iinibid．10(1)：60．1965——LamiumlongipetiolataHayata，Icon．Pl．Formos．8：92．1919．
本系概述：Ser．JavanicaeC．Y．WuetH．WLiinAct．Phytotax．Sin．10(1)：60．1965．萼齿明显，宽至狭三角形，先端钻形，刚毛状渐尖或具短尖头。
《Flora of China》 Vol. 17 (1994)
Paraphlomis javanica (Blume) Prain
假糙苏 jia cao su
Herbs erect. Stem solitary, 0.5-1.5 m, retrorse strigose, flexuous, leafless toward base. Petiole slender, to 8 cm; leaf blade elliptic to oblong-ovate, 3-15(-30) × 1.5-8.5(-14) cm, adaxially ± minutely hispid, abaxially strigose, base rounded to subtruncate, margin inconspicuously serrate to crenate-serrate, apex acute to acuminate. Verticillasters many flowered, globose, ca. 3 cm in diam., surrounded by few bracteoles at base; bracteoles subulate, ca. 6 mm, shorter than calyx, hirtellous. Flowers sessile. Calyx tubular in flower, abruptly spreading at mouth, enlarged and red in fruit, leathery, densely hirtellous or minutely hispid, glabrescent, inconspicuously veined, tube ca. 7 mm; teeth subequal, needlelike to triangular, 2-4 mm. Corolla yellow or yellowish, less often whitish, ca. 1.7 cm, ± hirtellous on apical parts of tube and limb outside, pubescent annulate inside, upper lip oblong, entire. Nutlets black, triquetrous, obovoid, apex rounded, glabrous. Fl. Jun-Aug, fr. Aug-Dec.
Evergreen and mixed forests; 300-2400 m. Fujian, Guangdong, Guangxi, Guizhou, Hainan, Hunan, Jiangxi, Sichuan, Taiwan, Yunnan [India, Indonesia, Laos, Malaysia, Myanmar, Pakistan, Philippines, Thailand, Vietnam].
