东亚羽节蕨Gymnocarpium oyamense (Bak.) Ching
东亚羽节蕨Gymnocarpium oyamense (Bak.) Ching
1.东亚羽节蕨(中国高等植物图鉴)大山羽节蕨(中国主要植物图说 蕨类植物门)、羽节蕨(台湾植物志)图版13:1-11
Gymnocarpium oyamense (Bak.) Ching in Contr. Biol. Lab. Sci. Soc. China Bot. Ser. 9: 40, f. 3. 1933; DeVol, Ferns East Centr. China in Notes Bot. Chin. Mus. Heude No. 7. 63, f. 31. 1945; 傅书遐, 中国主要植物图说 蕨类植物门 128,图 167. 1957; Tagawa, Col. Ill. Jap. Pterid. 114, 217, t. 44, f. 248. 1959; Namegata et Kurata, Enum. Jap. Pterid. 305, 1961; Ohwi, Fl. Jap. 75. 1965; Ic. Corm. Sin. 1: 173, f. 346. 1972; Fl. Tsinling. 2: 86. 1974; DeVol et C. M. Kuo in H. L. Li et al., Fl. Taiwan 1: 471. 1975; Nakaike, Enum. Pterid. Jap. 266. 1975 et New Fl, Jap. Pterid. 467, cum f. 1982 et l. c. Rev. Enl. 1992; Y. L. Chang et al., Sporae Pterid. Sin. 230, t. 40, f. 6-7. 1976; 河南植物志 1: 48, f. 60. 1981; Ching et Z. R. Wang in C. Y. Wu, Fl. Xizang. 1: 110. 1983; Z. R. Wang in W. T. Wang, Vasc. Pl. Hengduan Mount. 1: 65. 1993; J. F. Cheng in J. F. Cheng et G. F. Zhu, Fl. Jiangxi 1: 140, f. 121. 1993; W. C. C. F. Zhang in C. F. Zhang et S. Y. Zhang, Fl. Zhejiang 1: 114, f. 1-122. 1993; Shieh et al. in T. C. Huang Fl. Taiwan, 2nd ed. 1: 445. 1994; S. F. Wu in W. T. Wang, Keys Vasc. Pl. Wuling Mount. 34. 1995. ——Polypodium oyamense Bak. in Journ. Bot. 6: 366. 1877. ——Dryopteris oyamensis C. Chr., Ind. Fil. 282. 1906 et Suppl. 3: 94. 1934; Ogata, Ic. Fil. Jap. 1: t. 21. 1928. ——Currania oyamensis Copel. in Philip. Journ. Sci. Bot. 6: 147. 1909; Ohwi, Fl. Jap. Pterid. 101. 1957. ——Phegopteris oyamensis v. A. v. R., Mal. Ferns Suppl, 312. 1917. ——Carpogymnia oyamensis Love et Love in Taxon 16: 191. 1967. ——Polypodium krameri Franch. et Sav., Enum. Pl. Jap. 2: 244. 1876, nom. nud. et 2: 641. 1879. ——Nephrodium krameri Diels in Engl. u. Prantl, Nat. Pflanzenfam. 1 (4): 170. 1899. ——Phegopteris krameri Makino, Phan, et Pterid. Jap. Ic. Ill. 1: t. 30. 1899. ——Aspidium krameri Christ in Bull. Soc. France 52 Mem. 1: 37. 1905. ——Polypodium krameri var. incisum Franch. et Sav. Enum. Pl. Jap. 2: 244. 1877. ——Currania oyamensis Copel. var. incisa Koidz., Fl. Symb. Orient. -Asiat. 38. 1930. nom. nud. ——Polypodium gymnogrammoides Bak. in Journ. Bot. 26: 229. 1888. ——Nephrodium gymnogrammoides Diels in Engl. u. Prantl. Nat. Pflanzenfam. 1 (4): 170. 1899. ——Dryopteris gymnogrammoides C. Chr., Ind. Fil. 269. 1906. ——Currania gracilipes Copel. in Philip. Journ. Sci. Bot. 4: 112. 1909. ——Gymnocarpium gracilipes (Copel.) Ching in Contr. Biol. Lab. Sci. Soc. China Bot. Ser. 9: 39. 1933. ——Gymnocarpium oyamense var. gracilipes W. C. Shieh in Journ. Sci. Engin. (Nation. Chung-Hsing Univ.) 13: 50. 1976.
根状茎细长横走,直径约1.5-2(-3)毫米,被红褐色阔披针形鳞片,老时几光滑;叶远生。能育叶长 (13-) 25-45 (-50) 厘米;叶柄长 (7-) 12-25 (-31) 厘米,直径1.5-2毫米,禾秆色,有光泽,下面圆,上面有纵沟,基部被鳞片,向上几光滑,先端以关节和叶片相连;叶片卵状三角形,长 (7-) 10-18 (-22) 厘米,宽(4-) 6-13 (-20)厘米,先端渐尖,基部呈心形,一回羽状深裂达离叶轴4-5毫米处;裂片6-9 (-13)对,对生,平展,彼此接近,阔披针状镰刀形,先端向上弯,急尖或渐尖头,基部以阔翅彼此相连,边缘浅裂至深裂;基部一对裂片常为阔披针形,向下斜展,顶部弯向上,长(2-) 4-7 (-10)厘米,宽1-1.8 (-3) 厘米,第二对裂片较第一对略长,或几相等,平展,顶部也向上弯,从第三对裂片起向上逐渐变小,略斜向上;二回裂片有时不整齐,先端钝圆,边缘全缘或具浅圆齿。裂片上的主脉纤细,侧脉 (2-) 4-5 (-8)对,斜上,单一,略可见。叶干后草质,上面绿色,下面呈灰绿色,两面均无毛;叶轴基部与叶柄先端之间具有明显的关节,叶柄上部、叶轴下部和基部裂片的中脉基部生有稀疏的短腺体,或缺如。孢子囊群长圆形,生于裂片上的小脉中部,位于主脉两侧,彼此远离。孢子表面具有裂片状褶皱,上面具小穴状纹饰。染色体数目n=40,80。
分布于陕西(秦岭、长安县南五台、户县、佛坪、华县、华山、略阳、宁陕)、甘肃(康县)、安徽(黄山)、浙江(安吉、临安、遂昌、孝丰)、江西(庐山、井冈山、铜鼓、修水慕阜山)、台湾(高雄)、河南(嵩县、太行山、伏牛山、桐柏山、大别山)、湖北(宜昌)、湖南(桑植、石门、新宁)、四川(峨眉山、宝兴、天全)、重庆(南川)、贵州(毕节、纳雍、桐梓)、云南(丽江、大理、漾濞、禄劝、大关、永善)和西藏(察隅、南迦巴瓦)等省区。生林下湿地或石上苔藓中,海拔300-2900米。尼泊尔、日本、菲律宾和新几内亚岛也有分布。模式标本采自日本(神奈川县)。
《Flora of China》 Vol. 2-3 (2013)
Gymnocarpium oyamense (Baker) Ching Contrib. Biol. Lab. Sci. Soc. China, Bot. Ser. 9: 40. 1933.
东亚羽节蕨 dong ya yu jie jue
Polypodium oyamense Baker, J. Bot. 15: 366. 1877; Aspidium krameri (Franchet & Savatier) Christ; Carpogymnia oyamensis (Baker) á. L?ve & D. L?ve; Currania gracilipes Copeland; C. oyamensis (Baker) Copeland; C. oyamensis var. gracilipes (Copeland) Tagawa; Dryopteris gracilipes (Copeland) C. Christensen; D. gymnogrammoides (Baker) C. Christensen; D. oyamensis (Baker) C. Christensen; Gymnocarpium gracilipes (Copeland) Ching; G. oyamense var. gracilipes (Copeland) W. C. Shieh; Nephrodium gymnogrammoides (Baker) Diels; N. krameri (Franchet & Savatier) Diels; Phegopteris krameri (Franchet & Savatier) Makino; P. oyamensis (Baker) Alderwerelt; Polypodium gymnogrammoides Baker; P. krameri Franchet & Savatier; P. krameri var. incisum Franchet & Savatier.
Rhizomes long creeping, 1.5-2(-3) mm in diam., clothed with red-brown, broadly lanceolate scales, glabrate when old. Fronds distant; fertile fronds (13-)25-45(-50) cm; stipe stramineous, shiny, (7-)12-25(-31) cm, 1.5-2 mm in diam., terete abaxially and grooved adaxially, scaly at base, upward glabrate, articulate to lamina at apex; lamina ovate-triangular, (7-)10-18(-22) × (4-)6-13(-20) cm, herbaceous, glabrate, base cordate, pinnatipartite 4-5 mm to rachis, apex acuminate; pinnae 6-9(-13) pairs, opposite, spreading, approximate, broadly lanceolate, falcate, base decurrent to narrow rachis wing, shallowly lobed to pinnatipartite, apex acute or acuminate; basal pair of pinnae descending, often broadly lanceolate, (2-)4-7(-10) × 1-1.8(-3) cm; second pair slightly longer than basal pinnae or subequal, spreading, shallowly lobed; pinna lobes obtuse-rounded, entire or crenate; veins pinnate, lateral veins (2-)4 or 5(-8) pairs, simple, slightly visible; rachis inserted obliquely and articulate to stipe; upper part of stipe, lower part of rachis, and bases of costae in lower pinnae sparsely pubescent with short glands, or glabrate. Sori oblong, medial on veins, biseriate along both sides of costules, exindusiate. Spore wall surface rugate, foveolate. 2n = 80, 160.
Damp areas in forests, on moss-covered rocks; 300-2900 m. Anhui, Chongqing, Gansu, Guizhou, Henan, Hubei, Hunan, Jiangxi, Shaanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [NE India, Japan, Nepal, New Guinea, Philippines].
别名:羽节蕨;亚东羽节蕨;东亚羽介蕨;大山羽节蕨;
科名:蹄盖蕨科 Athyriaceae
属名:羽节蕨属 Gymnocarpium
1.东亚羽节蕨(中国高等植物图鉴)大山羽节蕨(中国主要植物图说 蕨类植物门)、羽节蕨(台湾植物志)图版13:1-11
Gymnocarpium oyamense (Bak.) Ching in Contr. Biol. Lab. Sci. Soc. China Bot. Ser. 9: 40, f. 3. 1933; DeVol, Ferns East Centr. China in Notes Bot. Chin. Mus. Heude No. 7. 63, f. 31. 1945; 傅书遐, 中国主要植物图说 蕨类植物门 128,图 167. 1957; Tagawa, Col. Ill. Jap. Pterid. 114, 217, t. 44, f. 248. 1959; Namegata et Kurata, Enum. Jap. Pterid. 305, 1961; Ohwi, Fl. Jap. 75. 1965; Ic. Corm. Sin. 1: 173, f. 346. 1972; Fl. Tsinling. 2: 86. 1974; DeVol et C. M. Kuo in H. L. Li et al., Fl. Taiwan 1: 471. 1975; Nakaike, Enum. Pterid. Jap. 266. 1975 et New Fl, Jap. Pterid. 467, cum f. 1982 et l. c. Rev. Enl. 1992; Y. L. Chang et al., Sporae Pterid. Sin. 230, t. 40, f. 6-7. 1976; 河南植物志 1: 48, f. 60. 1981; Ching et Z. R. Wang in C. Y. Wu, Fl. Xizang. 1: 110. 1983; Z. R. Wang in W. T. Wang, Vasc. Pl. Hengduan Mount. 1: 65. 1993; J. F. Cheng in J. F. Cheng et G. F. Zhu, Fl. Jiangxi 1: 140, f. 121. 1993; W. C. C. F. Zhang in C. F. Zhang et S. Y. Zhang, Fl. Zhejiang 1: 114, f. 1-122. 1993; Shieh et al. in T. C. Huang Fl. Taiwan, 2nd ed. 1: 445. 1994; S. F. Wu in W. T. Wang, Keys Vasc. Pl. Wuling Mount. 34. 1995. ——Polypodium oyamense Bak. in Journ. Bot. 6: 366. 1877. ——Dryopteris oyamensis C. Chr., Ind. Fil. 282. 1906 et Suppl. 3: 94. 1934; Ogata, Ic. Fil. Jap. 1: t. 21. 1928. ——Currania oyamensis Copel. in Philip. Journ. Sci. Bot. 6: 147. 1909; Ohwi, Fl. Jap. Pterid. 101. 1957. ——Phegopteris oyamensis v. A. v. R., Mal. Ferns Suppl, 312. 1917. ——Carpogymnia oyamensis Love et Love in Taxon 16: 191. 1967. ——Polypodium krameri Franch. et Sav., Enum. Pl. Jap. 2: 244. 1876, nom. nud. et 2: 641. 1879. ——Nephrodium krameri Diels in Engl. u. Prantl, Nat. Pflanzenfam. 1 (4): 170. 1899. ——Phegopteris krameri Makino, Phan, et Pterid. Jap. Ic. Ill. 1: t. 30. 1899. ——Aspidium krameri Christ in Bull. Soc. France 52 Mem. 1: 37. 1905. ——Polypodium krameri var. incisum Franch. et Sav. Enum. Pl. Jap. 2: 244. 1877. ——Currania oyamensis Copel. var. incisa Koidz., Fl. Symb. Orient. -Asiat. 38. 1930. nom. nud. ——Polypodium gymnogrammoides Bak. in Journ. Bot. 26: 229. 1888. ——Nephrodium gymnogrammoides Diels in Engl. u. Prantl. Nat. Pflanzenfam. 1 (4): 170. 1899. ——Dryopteris gymnogrammoides C. Chr., Ind. Fil. 269. 1906. ——Currania gracilipes Copel. in Philip. Journ. Sci. Bot. 4: 112. 1909. ——Gymnocarpium gracilipes (Copel.) Ching in Contr. Biol. Lab. Sci. Soc. China Bot. Ser. 9: 39. 1933. ——Gymnocarpium oyamense var. gracilipes W. C. Shieh in Journ. Sci. Engin. (Nation. Chung-Hsing Univ.) 13: 50. 1976.
根状茎细长横走,直径约1.5-2(-3)毫米,被红褐色阔披针形鳞片,老时几光滑;叶远生。能育叶长 (13-) 25-45 (-50) 厘米;叶柄长 (7-) 12-25 (-31) 厘米,直径1.5-2毫米,禾秆色,有光泽,下面圆,上面有纵沟,基部被鳞片,向上几光滑,先端以关节和叶片相连;叶片卵状三角形,长 (7-) 10-18 (-22) 厘米,宽(4-) 6-13 (-20)厘米,先端渐尖,基部呈心形,一回羽状深裂达离叶轴4-5毫米处;裂片6-9 (-13)对,对生,平展,彼此接近,阔披针状镰刀形,先端向上弯,急尖或渐尖头,基部以阔翅彼此相连,边缘浅裂至深裂;基部一对裂片常为阔披针形,向下斜展,顶部弯向上,长(2-) 4-7 (-10)厘米,宽1-1.8 (-3) 厘米,第二对裂片较第一对略长,或几相等,平展,顶部也向上弯,从第三对裂片起向上逐渐变小,略斜向上;二回裂片有时不整齐,先端钝圆,边缘全缘或具浅圆齿。裂片上的主脉纤细,侧脉 (2-) 4-5 (-8)对,斜上,单一,略可见。叶干后草质,上面绿色,下面呈灰绿色,两面均无毛;叶轴基部与叶柄先端之间具有明显的关节,叶柄上部、叶轴下部和基部裂片的中脉基部生有稀疏的短腺体,或缺如。孢子囊群长圆形,生于裂片上的小脉中部,位于主脉两侧,彼此远离。孢子表面具有裂片状褶皱,上面具小穴状纹饰。染色体数目n=40,80。
分布于陕西(秦岭、长安县南五台、户县、佛坪、华县、华山、略阳、宁陕)、甘肃(康县)、安徽(黄山)、浙江(安吉、临安、遂昌、孝丰)、江西(庐山、井冈山、铜鼓、修水慕阜山)、台湾(高雄)、河南(嵩县、太行山、伏牛山、桐柏山、大别山)、湖北(宜昌)、湖南(桑植、石门、新宁)、四川(峨眉山、宝兴、天全)、重庆(南川)、贵州(毕节、纳雍、桐梓)、云南(丽江、大理、漾濞、禄劝、大关、永善)和西藏(察隅、南迦巴瓦)等省区。生林下湿地或石上苔藓中,海拔300-2900米。尼泊尔、日本、菲律宾和新几内亚岛也有分布。模式标本采自日本(神奈川县)。
《Flora of China》 Vol. 2-3 (2013)
Gymnocarpium oyamense (Baker) Ching Contrib. Biol. Lab. Sci. Soc. China, Bot. Ser. 9: 40. 1933.
东亚羽节蕨 dong ya yu jie jue
Polypodium oyamense Baker, J. Bot. 15: 366. 1877; Aspidium krameri (Franchet & Savatier) Christ; Carpogymnia oyamensis (Baker) á. L?ve & D. L?ve; Currania gracilipes Copeland; C. oyamensis (Baker) Copeland; C. oyamensis var. gracilipes (Copeland) Tagawa; Dryopteris gracilipes (Copeland) C. Christensen; D. gymnogrammoides (Baker) C. Christensen; D. oyamensis (Baker) C. Christensen; Gymnocarpium gracilipes (Copeland) Ching; G. oyamense var. gracilipes (Copeland) W. C. Shieh; Nephrodium gymnogrammoides (Baker) Diels; N. krameri (Franchet & Savatier) Diels; Phegopteris krameri (Franchet & Savatier) Makino; P. oyamensis (Baker) Alderwerelt; Polypodium gymnogrammoides Baker; P. krameri Franchet & Savatier; P. krameri var. incisum Franchet & Savatier.
Rhizomes long creeping, 1.5-2(-3) mm in diam., clothed with red-brown, broadly lanceolate scales, glabrate when old. Fronds distant; fertile fronds (13-)25-45(-50) cm; stipe stramineous, shiny, (7-)12-25(-31) cm, 1.5-2 mm in diam., terete abaxially and grooved adaxially, scaly at base, upward glabrate, articulate to lamina at apex; lamina ovate-triangular, (7-)10-18(-22) × (4-)6-13(-20) cm, herbaceous, glabrate, base cordate, pinnatipartite 4-5 mm to rachis, apex acuminate; pinnae 6-9(-13) pairs, opposite, spreading, approximate, broadly lanceolate, falcate, base decurrent to narrow rachis wing, shallowly lobed to pinnatipartite, apex acute or acuminate; basal pair of pinnae descending, often broadly lanceolate, (2-)4-7(-10) × 1-1.8(-3) cm; second pair slightly longer than basal pinnae or subequal, spreading, shallowly lobed; pinna lobes obtuse-rounded, entire or crenate; veins pinnate, lateral veins (2-)4 or 5(-8) pairs, simple, slightly visible; rachis inserted obliquely and articulate to stipe; upper part of stipe, lower part of rachis, and bases of costae in lower pinnae sparsely pubescent with short glands, or glabrate. Sori oblong, medial on veins, biseriate along both sides of costules, exindusiate. Spore wall surface rugate, foveolate. 2n = 80, 160.
Damp areas in forests, on moss-covered rocks; 300-2900 m. Anhui, Chongqing, Gansu, Guizhou, Henan, Hubei, Hunan, Jiangxi, Shaanxi, Sichuan, Taiwan, Xizang, Yunnan, Zhejiang [NE India, Japan, Nepal, New Guinea, Philippines].