亮毛蕨Acystopteris japonica (Luerss.) Nakai

别名：毛冷蕨;中华亮毛蕨;

科名：蹄盖蕨科 Athyriaceae

属名：亮毛蕨属 Acystopteris

《中国植物志》第3(2)卷039页
  1. 亮毛蕨（中国高等植物图鉴）毛冷蕨（台湾植物志）、中华亮毛蕨（中国蕨类植物孢子形态）图版8：1-11
  Acystopteris japonica (Luerss.) Nakai in Bot. Mag. Tokyo 47: 180. 1933; Tagawa in Acta Phytotax. Geobot. 4: 51. 1935; H. Ito, Fil. Jap. Ill. t. 207. 1944; Ching in Sunyatsenia 5: 237. 1950; Ohwi, Fl, Jap. Pterid. 105. 1957; Tagawa, Col. Ill. Jap. Pterid. 118, 174, t. 46, f. 259. 1959: Namegata et Kurata, Enum. Jap. Pterid. 275. 1961. Ic. Corm. Sin. 1: 176, f. 351. 1972; Nakaike, Enum. Pterid. Jap. 127. 1975 et New Fl. Jap. Pterid. 248, cum f. a, b. 1982 et l. c. Rev. Enl. 1992; Y. L. Chang et al., Sporae Pterid. Sin. 192, t. 40, f. 20-21. 1976; Fl. Fujian. 1: 94, f. 85. 1982; Shing in J. F. Cheng et G. F. Zhu, Fl. Jiangxi 1: 138, f. 119. 1993; C. F. Zhang in C. F. Zhang et S. Y. Zhang, Fl. Zhejiang 1: 114, f. 1-123, 1993; S. F. Wu in W. T. Wang, Keys Vasc. Pl. Wuling Mount. 34. 1995. ——Cystopteris japonica Luerss. in Engl. Bot. Jahrb. 4: 363. 1883; Bak. in Ann. Bot. 5: 203. 1891; Matsum., Ind. Pl. Jap. 1: 301. 1904; C. Chr., Ind. Fil. 204, 1906; Makino et Nemoto, Fl. Jap. 1597. 1925; Ogata, Ic. Fil. Jap. 1: t. 18. 1928; Ohwi, Fl. Jap. Pterid. 105. 1957; Blasdell, Mem. Tarr. Bot. Club. 21 (4): 48. 1963; DeVol et C. M. Kuo in H. L. Li et al., Fl. Taiwan 1: 457. 1975; Shieh et al. in T. C. Huang, Fl. Taiwan, 2nd ed. 1: 430. 1994. ——Cystopteris chinensis Ching in Y. L. Chang et al., Sporae Pterid. Sin. 191, t. 40, f. 23-24. 1976. nom. nud.
  根状茎横走，直径2-4毫米，疏被黄褐色披针形薄鳞片；叶近生。能育叶长 (12-) 40-60 (-90)厘米；叶柄长 (6-)15-25 (-40) 厘米，直径 (1-) 2-3 (-4) 毫米，栗黑色或紫褐色，上面有一条纵沟，基部疏被鳞片，向上几光滑，略有光泽；叶片阔卵形至卵状长圆形，长 (6-) 20-35 (-50) 厘米，宽 (3.5-) 15-18(-40)厘米，短渐尖或渐尖头，基部圆心形，二至三回羽状；羽片10-15对左右，基部一对不缩短，长圆形或阔披针形，长(1:8-) 8-15 (-20)厘米，宽(1-) 3-6(-10) 厘米，先端渐尖，基部近平截，水平开展或斜展，几无柄或有极短柄，对生或近对生，距上面一对3-4厘米，一至二回羽状；小羽片10-24对，羽轴下侧的较上侧的略长，基部一对稍缩短，近对生或上先出，第二对起长圆形，长 (0.5-) 2.5-4 (-5) 厘米，宽 (3-) 6-10 (-15)厘米，钝头，无柄，互生，基部近对称，平截或阔楔形，紧靠羽轴，羽裂或羽状；裂片5-14对，平展或略斜展，长圆形，钝头而有锯齿，两侧往往锐裂；第二对羽片与基部一对近等长，同形，但较狭，向上羽片渐短，顶部羽片披针形，一回羽状或羽裂；叶脉在裂片上羽状，小脉单一，达于叶边锯齿。叶干后草质，绿色，两面沿叶脉和各回羽轴、叶轴以及叶柄均疏被透明节状长毛，羽轴上还多少有鳞片状毛混生，叶轴和叶柄同色，羽轴禾秆色或仅基部与叶轴同色。孢子囊小，圆形，着生于裂片基部上侧小脉背部，在主脉两侧各排成整齐的一列；囊群盖小，膜质，灰绿色，卵圆形，边缘有腺状疏睫毛，仅以基部一点着生于囊托，被成熟的孢子囊群向后推开、压于下面宛如下位，宿存。孢子豆形或长圆球形、黄色，不具周壁，表面具密集的直立而头部膨大的棒状突起。染色体数目n=42。
  分布于浙江（淳安、临安、龙泉、庆阳、遂昌、松阳）、江西（井冈山、玉山）、福建（福州）、台湾（台北、桃园、新竹、台中、南投、嘉义）、湖北（兴山）、湖南（桑植、石门、永顺、沅陵）、广西（龙胜、大苗山、九万山）、四川（峨眉山、洪雅、筠连、古蔺）、重庆（缙云山、南川金佛山）、贵州（赤水、梵净山、惠水、江口、雷公山、荔波、松桃）和云南（昆明、禄劝、镇雄、巧家、大关、绥江）等省区。生沟谷林下，海拔400-2800米。也分布于日本。模式标本采自日本（鹿儿岛县）。

《Flora of China》 Vol. 2-3 (2013)
Acystopteris japonica  (Luerssen) Nakai Bot. Mag. (Tokyo). 47: 180. 1933.
亮毛蕨   liang mao jue
Cystopteris japonica Luerssen in Engler, Bot. Jahrb. Syst. 4: 363. 1883.
Plants summer-green. Rhizome creeping, 2-4 mm in diam., with sparse yellowish brown, broadly lanceolate, thinly membranous scales. Fronds approximate; fertile frond (12-) 40-60(-90) cm; stipe chestnut-black or purple-brown, (6-) 15-25(-40) cm, (1-)2-3(-4) mm in diam., grooved adaxially, with sparse scales at base, upward glabrate, shiny; lamina 2- or 3-pinnate, broadly ovate to deltoid-ovate, (6-)20-35(-50) × (3.5-)15-18(-40) cm, not narrowed to base, herbaceous, base subcordate, apex shortly acuminate or acuminate; pinnae 10-15 pairs, opposite or subopposite, 3-4 cm apart, spreading or ascending, subsessile or very shortly stalked, basal pair not shorter, oblong or broadly lanceolate; upper pinnae lanceolate, (1.8-)8-15(-25) × (1-)3-6(-11) cm, base subtruncate, apex acuminate; pinnules 10-24 pairs, sessile, alternate, subequilateral, basiscopic pinnules slightly longer than acroscopic pinnules; basal pair slightly shorter, subopposite or anadromous, oblong, (0.5-)2.5-4(-6) × (3-)6-10(-20) mm, base truncate or broadly cuneate, apex obtuse; costae chestnut-brown or stramineous in distal half; pinnule segments 5-14 pairs, spreading or slightly ascending, oblong, crenate, apex obtuse; veins pinnate in segment, simple, reaching marginal teeth. Hyaline multicellular long hairs and scalelike hairs sparse on both sides of veins, costae, rachis, and stipe. Sori orbicular, abaxial on basal acroscopic veins; indusia pale green, broadly ovate, small, membranous, glandular, sparsely ciliate at margin, proximal to sorophore, hidden by sporangia at maturity, persistent. Spores yellow, bean-shaped or long spheroidal, perispore with dense inflated, bulbous processes. 2n = 84*, 168*.
Forests in valleys; 400-2800 m. Chongqing, Fujian, Guangxi, Guizhou, Hubei, Hunan, Jiangxi, Yunnan, Zhejiang [Japan].